| 1965 - 1970 | Agriculture University, Prague (Diploma Thesis ”Polymorphism of transferrins in blood serum of Coregonus lavaretus maraena and Salvelinus fontinalis”). |
| 1971 - 1975 | study stay, Institute of Animal Physiology and Genetics, v.v.i., CZAS, Libechov |
| 1976 - 1980 | Ph.D. study, Institute of Animal Physiology and Genetics, v.v.i., CZAS, Libechov (Ph.D. Thesis ”Study of polymorphic protein systems in freshwater fishes”) |
| 1979 - 1985 | research assistant |
| 1986 - present | researcher |